Insight into tube-building behaviour and palaeoecology of some agglutinating worms from the Upper Devonian of Nevada, USA

Agglutinated worm tubes from the Upper Devonian of the Devils Gate section in Nevada, USA are reported for the first time, filling a major gap in their Palaeozoic fossil record. Two small (5 mm and 6.7 mm in length) tubes are composed entirely of tentaculitid shells, and one large tube (55 mm in length) is formed from particles including ostracode carapaces, echinoderm ossicles, tentaculitid shells and putative bryozoan fragments aligned perpendicularly to the tube’s long axis. The tubes, in particular the large one have a cylindrical, curved and tapering tube morphology that is very similar to that of modern agglutinating polychaetes of the families Terebellidae and Pectinariidae. The large tube is dominated by objects that fall within a certain size-range, and although built from different types of particles, echinoderm ossicles are prevalent in the posterior part, whereas ostracode carapaces dominate in the middle and anterior parts of the tube. Tentaculitid shells are relatively rare in the large tube, despite being abundant in the surrounding host deposit. The faunal assemblage composing the tube suggests that the worm animal was rather specific in its selection of particles with a certain morphology. This is common behaviour amongst many modern agglutinating terebellid and pectinariid polychaetes. The preservation of such fragile tubes was enhanced by rapid burial, likely caused by gravity flow of sediment in a deep-slope setting

Large igneous provinces and mass extinctions: an update

The temporal link between mass extinctions and large igneous provinces is well known. Here, we examine this link by focusing on the potential climatic effects of large igneous province eruptions during several extinction crises that show the best correlation with mass volcanism: the Frasnian-Famennian (Late Devonian), Capitanian (Middle Permian), end-Permian, end-Triassic, and Toarcian (Early Jurassic) extinctions. It is clear that there is no direct correlation between total volume of lava and extinction magnitude because there is always sufficient recovery time between individual eruptions to negate any cumulative effect of successive flood basalt eruptions. Instead, the environmental and climatic damage must be attributed to single-pulse gas effusions. It is notable that the best-constrained examples of death-by-volcanism record the main extinction pulse at the onset of (often explosive) volcanism (e.g., the Capitanian, end-Permian, and end-Triassic examples), suggesting that the rapid injection of vast quantities of volcanic gas (CO 2 and SO 2 ) is the trigger for a truly major biotic catastrophe. Warming and marine anoxia feature in many extinction scenarios, indicating that the ability of a large igneous province to induce these proximal killers (from CO 2 emissions and thermogenic greenhouse gases) is the single most important factor governing its lethality. Intriguingly, many voluminous large igneous province eruptions, especially those of the Cretaceous oceanic plateaus, are not associated with significant extinction losses. This suggests that the link between the two phenomena may be controlled by a range of factors, including continental configuration, the latitude, volume, rate, and duration of eruption, its style and setting (continental vs. oceanic), the preexisting climate state, and the resilience of the extant biota to change

Ultra-shallow-marine anoxia in an Early Triassic shallow-marine clastic ramp (Spitsbergen) and the suppression of benthic radiation

Lower Triassic marine strata in Spitsbergen accumulated on a mid-to-high latitude ramp in which high-energy foreshore and shoreface facies passed offshore into sheet sandstones of probable hyperpycnite origin. More distal facies include siltstones, shales and dolomitic limestones. Carbon isotope chemostratigraphy comparison allows improved age dating of the Boreal sections and shows a significant hiatus in the upper Spathian. Two major deepening events, in earliest Griesbachian and late Smithian time, are separated by shallowing-upwards trends that culminated in the Dienerian and Spathian substages. The redox record, revealed by changes in bioturbation, palaeoecology, pyrite framboid content and trace metal concentrations, shows anoxic phases alternating with intervals of better ventilation. Only Dienerian–early Smithian time witnessed persistent oxygenation that was sufficient to support a diverse benthic community. The most intensely anoxic, usually euxinic, conditions are best developed in offshore settings, but at times euxinia also developed in upper offshore settings where it is even recorded in hyperpycnite and storm-origin sandstone beds: an extraordinary facet of Spitsbergen's record. The euxinic phases do not track relative water depth changes. For example, the continuous shallowing upwards from the Griesbachian to lower Dienerian was witness to several euxinic phases separated by intervals of more oxic, bioturbated sediments. It is likely that the euxinia was controlled by climatic oscillations rather than intra-basinal factors. It remains to be seen if all the anoxic phases found in Spitsbergen are seen elsewhere, although the wide spread of anoxic facies in the Smithian/Spathian boundary interval is clearly a global event

Lower Wenlock black shales in the northern Holy Cross Mountains, Poland: Sedimentary and geochemical controls on the Ireviken Event in a deep marine setting

The stratigraphic variability and geochemistry of Llandovery/Wenlock (L/W) Series boundary sediments in Poland reveals that hemipelagic sedimentation under an anoxic/euxinic water column was interrupted by low density bottom currents or detached diluted turbid layers that resulted in intermittent seafloor oxygenation. TOC values and inorganic proxies throughout the Wilków 1 borehole section suggest variable redox conditions. U/Mo ratios >1 throughout much of the Aeronian and Telychian Stages, together with an absence of pyrite framboids, suggests oxygenated conditions prevailed. However, elevated TOC near the Aeronian/Telychian boundary, together with increased U/Th and V/(V+Ni) ratios and populations of small pyrite framboids are consistent with the development of dysoxic/anoxic conditions at that time. U/Th, V/Cr and V/(V+Ni) ratios, as well as Uauthig and Mo concentrations suggest that during the Ireviken black shale (IBS) deposition, bottom-water conditions deteriorated from oxic during the Telychian to mostly suboxic/anoxic immediately prior to the L/W boundary, before a brief reoxygenation at the end of the IBS sedimentation in the Sheinwoodian Stage. Rapid fluctuations in U/Mo during the Ireviken Event (IE) are characteristic of fluctuating redox conditions that culminated in an anoxic/euxinic seafloor in the Sheinwoodian. Following IBS deposition, conditions once again became oxygen deficient with the development of a euxinic zone in the water column. The Aeronian to Sheinwoodian deep-water redox history was unstable, and rapid fluctuations of the chemocline across the L/W Series boundary probably contributed to the IE extinctions, which affected mainly pelagic and hemipelagic fauna

The great catastrophe: causes of the Permo-Triassic marine mass extinction

The marine losses during the Permo-Triassic mass extinction were the worst ever experienced. All groups were badly affected, especially amongst the benthos (e.g. brachiopods, corals, bryozoans, foraminifers, ostracods). Planktonic populations underwent a fundamental change with eukaryotic algae being replaced by nitrogen-fixing bacteria, green-sulphur bacteria, sulphate-reducing bacteria and prasinophytes. Detailed studies of boundary sections, especially those in South China, have resolved the crisis to a ∼55 kyr interval straddling the Permo-Triassic boundary. Many of the losses occur at the beginning and end of this interval painting a picture of a two-phase extinction. Improved knowledge of the extinction has been supported by numerous geochemical studies that allow diverse proposed extinction mechanisms to be studied. A transition from oxygenated to anoxic-euxinic conditions is seen in most sections globally, although the intensity and timing shows regional variability. Decreased ocean ventilation coincides with rapidly rising temperatures and many extinction scenarios attribute the losses to both anoxia and high temperatures. Other kill mechanisms include ocean acidification for which there is conflicting support from geochemical proxies and, even less likely, siltation (burial under a massive influx of terrigenous sediment) which lacks substantive sedimentological evidence. The ultimate driver of the catastrophic changes at the end of the Permian was likely Siberian Trap eruptions and their associated carbon dioxide emissions with consequences such as warming, ocean stagnation and acidification. Volcanic winter episodes stemming from Siberian volcanism have also been linked to the crisis, but the short-term nature of these episodes (<decades) and the overwhelming evidence for rapid warming during the crisis makes this an unlikely cause. Finally, whilst the extinction is well studied in equatorial latitudes, a different history is found in northern Boreal latitudes including an earlier crisis which merits further study in order to fully understand the course and cause of the Permo-Triassic extinctions

An abrupt extinction in the Middle Permian (Capitanian) of the Boreal Realm (Spitsbergen) and its link to anoxia and acidification

The controversial Capitanian (Middle Permian, 262 Ma) extinction event is only known from equatorial latitudes, and consequently its global extent is poorly resolved. We demonstrate that there were two, severe extinctions amongst brachiopods in northern Boreal latitudes (Spitsbergen) in the Middle to Late Permian, separated by a recovery phase. New age dating of the Spitsbergen strata (belonging to the Kapp Starostin Formation), using strontium isotopes and d13C trends and comparison with better-dated sections in Greenland, suggests that the first crisis occurred in the Capitanian. This age assignment indicates that this Middle Permian extinction is manifested at higher latitudes. Redox proxies (pyrite framboids and trace metals) show that the Boreal crisis coincided with an intensification of oxygen depletion, implicating anoxia in the extinction scenario. The widespread and near-total loss of carbonates across the Boreal Realm also suggests a role for acidification in the crisis. The recovery interval saw the appearance of new brachiopod and bivalve taxa alongside survivors, and an increased mollusk dominance, resulting in an assemblage reminiscent of younger Mesozoic assemblages. The subsequent end-Permian mass extinction terminated this Late Permian radiation

Present and future evidence for evolving dark energy

We compute the Bayesian evidences for one- and two-parameter models of evolving dark energy, and compare them to the evidence for a cosmological constant, using current data from Type Ia supernova, baryon acoustic oscillations, and the cosmic microwave background. We use only distance information, ignoring dark energy perturbations. We find that, under various priors on the dark energy parameters, LambdaCDM is currently favoured as compared to the dark energy models. We consider the parameter constraints that arise under Bayesian model averaging, and discuss the implication of our results for future dark energy projects seeking to detect dark energy evolution. The model selection approach complements and extends the figure-of-merit approach of the Dark Energy Task Force in assessing future experiments, and suggests a significantly-modified interpretation of that statistic.Comment: 10 pages RevTex4, 3 figures included. Minor changes to match version accepted by PR

The Canada-UK Deep Submillimetre Survey: First Submillimetre Images, the Source Counts, and Resolution of the Background

We present the first results of a deep unbiased submillimetre survey carried out at 450 and 850 microns. We detected 12 sources at 850 microns, giving a surface density of sources with 850-micron flux densities > 2.8mJy of of 0.49+-0.16 per square arcmin. The sources constitute 20-30% of the background radiation at 850 microns and thus a significant fraction of the entire background radiation produced by stars. This implies, through the connection between metallicity and background radiation, that a significant fraction of all the stars that have ever been formed were formed in objects like those detected here. The combination of their large contribution to the background radiation and their extreme bolometric luminosities make these objects excellent candidates for being proto-ellipticals. Optical astronomers have recently shown that the UV-luminosity density of the universe increases by a factor of about 10 between z=0 and z=1 and then decreases again at higher redshifts. Using the results of a parallel submillimetre survey of the local universe, we show that both the submillimetre source density and background can be explained if the submillimetre luminosity density evolves in a similar way to the UV-luminosity density. Thus, if these sources are ellipticals in the process of formation, they may be forming at relatively modest redshifts.Comment: 8 pages (LATEX), 6 postscript figures, submitted to ApJ Letter

Circles in the Sky: Finding Topology with the Microwave Background Radiation

If the universe is finite and smaller than the distance to the surface of last scatter, then the signature of the topology of the universe is writ large on the microwave background sky. We show that the microwave background will be identified at the intersections of the surface of last scattering as seen by different ``copies'' of the observer. Since the surface of last scattering is a two-sphere, these intersections will be circles, regardless of the background geometry or topology. We therefore propose a statistic that is sensitive to all small, locally homogeneous topologies. Here, small means that the distance to the surface of last scatter is smaller than the ``topology scale'' of the universe.Comment: 14 pages, 10 figures, IOP format. This paper is a direct descendant of gr-qc/9602039. To appear in a special proceedings issue of Class. Quant. Grav. covering the Cleveland Topology & Cosmology Worksho

Correlated adiabatic and isocurvature perturbations from double inflation

It is shown that double inflation (two minimally coupled massive scalar fields) can produce correlated adiabatic and isocurvature primordial perturbations. Depending on the two relevant parameters of the model, the contributions to the primordial perturbations are computed, with special emphasis on the correlation, which can be quantitatively represented by a correlation spectrum. Finally the primordial spectra are evolved numerically to obtain the CMBR anisotropy multipole expectation values. It turns out that the existence of mixing and correlation can alter very significantly the temperature fluctuation predictions.Comment: 23 pages, 7 figures, RevTex. To appear in Phys. Rev.